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Tawny nurse shark
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Tawny nurse shark : ウィキペディア英語版
Tawny nurse shark

The tawny nurse shark (''Nebrius ferrugineus'') is a species of carpet shark in the family Ginglymostomatidae, and the only extant member of the genus ''Nebrius''. It is found widely along coastlines in the Indo-Pacific, preferring reefs, sandy flats, and seagrass beds from very shallow water to a depth of . With a cylindrical body and a broad, flattened head, the tawny nurse shark is quite similar in appearance to the nurse shark (''Ginglymostoma cirratum'') of the Atlantic and East Pacific, from which it can be distinguished by its pointed-tipped dorsal fins and narrow, sickle-shaped pectoral fins. The maximum recorded length of the tawny nurse shark is .
Nocturnal in habits, the tawny nurse shark tends to spend the day resting in piles of two dozen or more individuals inside caves or under ledges. At night, it is an active-swimming predator that uses a powerful suction force to extract prey from inside holes and crevices. The diet of this species consists mainly of octopus, though they also take other invertebrates, small bony fishes, and rarely sea snakes. It is aplacental viviparous, meaning the embryos hatch from egg capsules inside the mother. It is the only carpet shark in which the embryos are oophagous, feeding on eggs produced by the mother while inside the uterus. The litter size may be as small as one or two, based on the large size of near-term embryos.
Compared to the nurse shark, the tawny nurse shark has a more placid disposition and will often allow divers to touch and play with it. However, it should be accorded respect due to its powerful jaws and sharp teeth. This species is caught by commercial fisheries across most of its range for meat, fins, liver oil, leather, and fishmeal. It is also esteemed as a game fish off Queensland, Australia, and is known for its habit of spitting water in the faces of its captors. The International Union for Conservation of Nature (IUCN) has assessed the tawny nurse shark as Vulnerable, with subpopulations in several areas already diminished or extirpated.
==Taxonomy and phylogeny==

The tawny nurse shark was first described by French naturalist René-Primevère Lesson as ''Scyllium ferrugineum'', based on a long specimen from New Guinea. His short account was published in 1831 in ''Voyage au tour du monde, sur la corvette La Coquille''. A more detailed description, along with an illustration, was published by German naturalist Eduard Rüppell in 1837 as ''Nebrius concolor'', based on a specimen from the Red Sea. Both names were retained, often in separate genera (''Ginglymostoma'' and ''Nebrius'' respectively), until they were synonymized by Leonard Compagno in 1984. Compagno recognized that the tooth shape differences used to separate these species were the result of differences in age, with ''N. concolor'' representing younger individuals.
The genus name ''Nebrius'' is derived from the Greek word ''nebris'' or ''nebridos'', meaning the skin of a fawn. The specific epithet ''ferrugineus'' is Latin for "rust-colored".〔Bester, C. (Biological Profiles: Tawny Nurse Shark ). Florida Museum of Natural History Ichthyology Department. Retrieved on June 7, 2009.〕 Other common names for this species include giant sleepy shark, Madame X (a name coined by the shark fisherman Norman Caldwell in the 1930s for the then-unidentified Australian specimens), nurse shark, rusty catshark, rusty shark, sleepy shark, spitting shark, and tawny shark. Based on morphological similarities, ''Nebrius'' is believed to be the sister genus of ''Ginglymostoma'', with both being placed in a clade that also contains the short-tail nurse shark (''Pseudoginglymostoma brevicaudatum''), the whale shark (''Rhincodon typus''), and the zebra shark (''Stegostoma fasciatum'').

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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